Major General Issam Zahreddine, who led his Syrian Republican Guard brigade for three years against an ISIS siege in Deir Ezzor, was reportedly killed by a landmine.
"Lie in wait for them at every chance to ambush. The Levant is the graveyard of the Russians and Americans," reads text on a recent propaganda image.
Syria's war is almost over and experts say the U.S. has few options to counter what's been a major victory for Russia, Iran and President Bashar al-Assad.
The accusation is the latest sign of rising tensions between Moscow and Washington.
The Syrian military's Fifth Assault Corps, known as "ISIS Hunters," has made ISIS an offer it can't refuse.
The U.S. denies Moscow's allegations, as the Kremlin has a history of conflating the "terrorist" groups it claims to fight in Syria.
In the video, a man identifies himself and a companion as Russian soldiers captured by ISIS during a counteroffensive against Syrian troops in Deir Ezzor.
The 2,000-year-old Lion of al-Lat statue is a symbol of peace, but its restoration is only a reprieve from the death and destruction Palmyra has witnessed.
Russian President Vladimir Putin said "the necessary conditions have been created for putting an end to the fratricidal war in Syria."
The most wanted extremist in the world had not spoken since November 2016.
The Iraqi prime minister rejected the Kurdish vote for independence and parliament has given him military power to oppose it.
Turkey and Iran, traditional foes, have established common ground in Iraq, Syria and Qatar as regional alliances shift.
The British men included Jihadi John, executioner of three Americans, and Junaid Hussain, ISIS's cyber chief.
Moscow is sending an orchestra for Greece and cruise missiles for Syria in one go, according to reports.
Two rivals are battling ISIS in the jihadists' final stronghold of Deir ez-Zor. One seeks to return Syria to its former self; the other wants more power for Kurds.
Nicaragua says it will sign up to the climate deal, leaving Trump and Assad the only ones outside.
Russian and U.S. allies are battling insurgents and jihadists trying to topple the Syrian government, but the latter wants to reshape the nation.
The U.S. has designated the Al-Qaeda affiliated a "terrorist" group, despite its attempts to rebrand. However Russia has repeatedly accused Washington of sparing its militants.
The jihadist group is cranking up its official output as its ranks and morale drop.
The Kentucky senator threatens to try to delay passage of the national defense bill if there isn't a debate on his amendment.
German investigators create list of serial numbers that could be used to provide fake identities for jihadists.
Israel delivered two chickens a day for the eagle to eat, while the troops faced a food shortage.