The operation seized thousands of animals and identified around 1,400 people suspected of related criminal activity.
The radiated tortoise is a popular pet in Asia, and some smuggle them out of Madagascar to meet demand.
He reportedly has run his illegal business for more than a decade.
One account said this was the worst animal welfare violation she'd ever seen.
Wildlife trafficking continues to affect the elephants, giraffes, buffalo and antelope in Garamba.